Latitudinal range: 38.3° to 41.6°

Found in 4 samples

Bulgaria: Gorna Sushitsa (Late Miocene)

Greece: Halmyropotamos (Late Miocene)

Turkey: Kemiklitepe (KTA) (Late Miocene), Kemiklitepe (KTB) (Late Miocene)

See also Skoufotragus